小陇山锐齿栎林种群空间格局及关联性变化分析
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引用本文:马映栋,刘文桢,赵中华,石小龙,李安明.小陇山锐齿栎林种群空间格局及关联性变化分析[J].西北植物学报,2014,34(9):1878~1886
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作者单位
马映栋1,刘文桢2*,赵中华3,石小龙2,李安明2 (1 甘肃农业大学 林学院,兰州 730700
2 甘肃小陇山林业科学研究所,甘肃天水 741020
3 中国林业科学院 林业研究所,北京 100049) 
基金项目:“十二五”国家科技支撑计划课题(2012BAD22B03)
中文摘要:基于2008年和2013年对位于秦岭山脉西端、甘肃东南部小陇山林区的林分调查数据,选取树种锐齿栎、膀胱果和青榨槭为研究对象,运用点格局O-ring O(t)函数分析法,探讨5年前后锐齿栎林种群空间分布格局与关联性变化情况(其中,对树种锐齿栎进行分林层讨论,而对于膀胱果和青榨槭则不分层,空间关联性分析均不分层)。结果显示:(1)5年前后,锐齿栎林的空间格局整体上表现为随机分布;林上层锐齿栎分布稳定,主要为随机分布,林间层多均匀分布,林下层聚集分布明显;树种膀胱果和青榨槭因株数较少,在大部分尺度上仍随机分布,但随时间变化在局部尺度上则出现轻微的聚集分布。(2)分析林木空间关联性变化,则膀胱果与锐齿栎在幼树期因相互竞争而大部分尺度上互为负相关,其生长发育到一定阶段相互促生且小部分尺度上相互独立;树种锐齿栎与青榨槭、膀胱果与青榨槭在整体上保持关联,逐渐发展为小范围内互为相关性。研究表明:不同调查时间锐齿栎林主要树种空间分布格局变化对其尺度依赖性强,林木分布格局的尺度吻合程度较高,林分整体上为随机分布,说明五年前后林分空间分布格局基本没变;就主要树种而言,锐齿栎分布格局变化符合群落中种群发育的规律,膀胱果和青榨槭分布格局变化受株数影响较大;林木种间关联性变化与种群结构及其演替阶段有关;但就群落演替过程而言,5年期限不足以表征整座林分的演替规律,需进一步拓宽研究年限。
中文关键词:林分  尺度  空间分布格局  关联性  变化  小陇山
 
Change Analysis of Spatial Pattern and Correlation for Quercus aliena var.acuteserrata Forest Population in Xiaolong Mountains
Abstract:Based on data information investigated twice in Xiaolongshan forest region of west of Qinling Mountains in 2008 and 2013,we chose trees Quercus aliena var.acuteserrata,Staphylea holocarpa and Acer davidii as research objects,and then changes of spatial pattern and correlation for the above trees in five years were analyzed with O-ring O(t) method of point pattern.Spatial pattern of Q.aliena var.acuteserrata must be discussed depending on different forest layers,but that of S.holocarpa and A.davidii and correlation of all population would not rest on forest layers for further study.The results show that:(1)Spatial pattern distribution of forest stand Q.aliena var.acuteserrata was,as a whole,random over five years.Q.aliena var.acuteserrata distributed randomly in the upper forest layer,more evenly in the middle and clumped obviously in the lower.The small number of S.holocarpa and A.davidii led to them with random distribution on most scales and then later,along with time variation,to them with lightly aggregated distribution on small part of scales.(2)For changes of spatial population correlation,there was negative correlation between S.holocarpa and Q.aliena var.acuteserrata on most scales for competition at the stage of sapling,and independent distribution on small part of scales to the certain stage of growth.The relation between Q.aliena var.acuteserrata and A.davidii,and S.holocarpa and A.davidii kept independent on the whole,but gradually the correlation existed on a small scale.The research indicates that,in different survey time,change of spatial distribution pattern for dominant trees in Q.aliena var.acuteserrata forest depended strongly on the scale,goodness of fit of the distribution pattern was high,and the forest stand on the whole was random distribution,namely spatial distribution pattern remained largely unchanged in five years;for principal species,the change of distribution pattern of Q.aliena var.acuteserrata conformed to growth regulation of communities and that of S.holocarpa and A.davidii was more affected by number of trees;change of forest spatial correlation was connected with population structure and its stage of succession;but for successional process of community,successional change of forest community in five years won’t enough represent successional rule of all forest stand,and the study must be still carried out over a long period.
keywords:forest stand  scale  spatial distribution pattern  correlation  change  Xiaolong Mountains
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