林分密度对桉树人工林群落结构和物种多样性的影响
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国家自然科学基金(31370628);


Effects of Stand Density on Community Structure and Species Diversity of Eucalyptus robusta Plantation
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    摘要:

    采用典型样地法,以四川新津县普兴镇桉树(Eucalyptus robusta)人工林为研究对象,设置5种不同林分密度(A. 625 株·hm-2;B. 750 株·hm-2;C. 875 株·hm-2;D. 1 000 株·hm-2;E. 1 125 株·hm-2)样地各3个,对不同林分密度人工林群落结构和物种多样进行分析,探讨确定桉树人工林的合理栽植密度,为维持桉树人工林群落结构稳定、增加物种多样性提供依据。结果表明:(1)研究区桉树人工林群落共记录到96种植物,隶属于51科79属;在各个林分密度的群落中,植物科属种数量变化规律均为:草本层>灌木层>乔木层;当密度为1 000 株·hm-2时,乔木层物种数最高(6种);当密度为750、 875 株·hm-2时,灌木层群落物种数最高(17种);在林分密度625 株·hm-2下,草本层物种数最高(32种)。(2)桉树人工林群落各层次的丰富度指数D值、ShannnonWiener指数H值、Simpson指数H′、Pielou均匀度指数Jsw值均表现为:草本层>灌木层>乔木层;乔木层的D值和H′值在875 株·hm-2密度下最大,H值在625 株·hm-2密度下最大,Jsw值在750 株·hm-2密度下最大;灌、草层各指数分别均在密度750、625 株·hm-2时有最大值。(3)从群落结构上看,5种林分密度的群落径级结构均呈单峰型分布,乔木个体多分布在中径级(7.0≤d<23.0 cm);高度级结构与径级结构对林分密度的响应不一致,密度A、B、C和D、E的个体分别集中在高度级Ⅱ~Ⅴ(6.0≤h<18.0 m)、Ⅲ~Ⅴ(9.0≤h<18.0 m)、Ⅱ~Ⅲ(6.0≤h<12.0 m)和Ⅳ~Ⅴ(12.0≤h<18.0 m)。(4)各层次优势种不同,乔木层中桉树是绝对优势树种,各密度林分中构树(Broussonetia papyrifera)的重要值基本仅次于桉树;灌木层中,桉树幼苗只在密度E出现,构树在各密度下优势地位显著,在密度B~D中,均有八角枫(Alangium chinense)和女贞(Ligustrum lucidum)存在,并在密度B下八角枫(0.118 4)和女贞(0.183 2)长势较好,具有较大的更新潜力;青蒿(Artemisia carvifolia)和荩草(Arthraxon hispidus)作为草本层的优势种在5种林分密度下都有生长。研究认为,林分密度750株·hm-2(密度B)是新津桉树人工林的最适林分密度,该密度更有利于该地区桉树人工林群落结构和物种多样性的稳定可持续发展。

    Abstract:

    In order to find a reasonable density to maintain a stable community structure and improve species diversity, we studied the effects of different stand densities on community structure and species diversity of Eucalyptus robusta plantation in Puxing Town, Xinjin County, Sichuan Province, using typical sampling method, setting 3 plots in each stand density (A. 625 trees·hm-2; B. 750 trees·hm-2; C. 875 trees·hm-2; D. 1 000 trees·hm-2; E. 1 125 trees·hm-2). The results showed that: (1) a total of 96 plants were recorded, belonging to 51 families and 79 genera. The number of species in the vegetation of every kind of stand density arranged from greatest to least was: the herb layer, the shrub layer, and the tree layer. When the density was 1 000 trees·hm-2, there were 6 species in the tree layer, which was the maximum. And the maximum was 17 species in the shrub layer when the densities were 750 and 875 trees·hm-2 , there were 32 species in the herb layer under the stand density 625 trees·hm-2.(2) In general, the species richness index D, ShannnonWiener index H, Simpson index H′ and Pielou uniformity index Jsw of the E. robusta plantation community in all kinds of stand densities were all expressed as: herb layer>shrub layer>tree layer. The maximum of the D and H′, H, Jsw value of the tree layer was respectively from the density C, A, B; each index of the shrub and herb layer had the maximums respectively at the density 750 and 625 trees·hm-2. (3) Seen from the community structure, the diameter class of the five kinds of stand densities showed a single peak type distribution, and the trees were mostly distributed in the middle diameter classes(7.0≤d<23.0 cm); the height class was different from the diameter class, the trees in the density of A, B, C and D, E were concentrated respectively in the height classes Ⅱ to Ⅴ(6.0≤h<18.0 m), Ⅲ to Ⅴ(9.0≤h<18.0 m), Ⅱ to Ⅲ(6.0≤h<12.0 m), Ⅳ to Ⅴ(12.0≤h<18.0 m). (4) The dominant tree species varied among different layers. It was E. robusta in the tree layer, and the important value of Broussonetia papyrifera in each density was second only to E. robusta in general. In the shrub layer, E. robusta seedlings only appeared in density E, and the dominant position of B. papyrifera in each density was significant. In density B-D, both Alangium chinense and Ligustrum lucidum existed, in which they grew well with the important value of 0.118 4 and 0.183 2 under density B, having great potential for renewal. As the dominant species of herb layer, Artemisia carvifolia and Arthraxon hispidus both grew under the five kinds of stand densities. The results showed that the stand density 750 trees·hm-2 was the optimum for the E. robusta plantation, which was more conducive to the stable and sustainable development of community structure and species diversity of E. robusta plantation in this area.

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张柳桦,齐锦秋,柳苹玉,等.林分密度对桉树人工林群落结构和物种多样性的影响[J].西北植物学报,2018,38(1):166-175

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  • 在线发布日期: 2018-02-05
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