Abstract:The megasporogenesis, microsporogenesis and development of female and male gametophyte of Tupistra chinensis Baker were observed by the method of paraffin sections. The main results are follows: (1)the anther of T. chinensis Baker is four sporangiate and the development of anther wall is the basic type, and the anther wall is composed of epidermis, endothecium, middle layer and tapetum. The epidermis consisits of one layer of cells and still existed as the anther matured. The endothecium developed bandingshaped thickenings when the anther matured. The young anther wall has 2-3 layer middle layers and degenerated when maturity of the anther. The tapetum belongs to secretory tapetum and degenerated at the time of anther matured. (2)The cytokinesis of the microspore mother cells in meiosis is of successive type. The microspore tetrads are isobilateral spore. The mature pollen grain is of the 2celled type with single germinal furrow.(3)The ovary has three chamber which has 6 ovules that is double integuments, tenuinucellate and anatropous. The archesporial cell divides transversely, cutting an outer parietal cell and an inner sporogenous cell. The development of the embryo sac belongs to bisporic embryo scas. After the first meiosis, the megasporocyte divides into dyad cells and only the chalazal dyad cell undergoes the second meiotic division, whereas the micropylar dyad cell degenerates. Both the megaspore nuclei which from the functional dyad cell contribute in the formation of embryo sac. The embryo sac is of a Allum type.